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DARK AGES

DARMSTADT

681

yield in 1883 was 8,080,293 lb. See the Report of off by nine o'clock A.M. Such an undertaking, so the Indian Survey for 1883-84.

Dark Ages. See AGE.

Darley, FELIX O. C., an eminent American artist, was born at Philadelphia, June 23, 1822. He early gave himself to drawing with such success as to encourage him to devote himself exclusively to art, especially in the form of book illustration. His earliest important work was a series of drawings for the Library of Humorous American Works, after producing which he went to New York (1848), where his outline drawings to Washington Irving's Legend of Sleepy Hollow and Rip Van Winkle soon spread wide his reputation. Later works, not less popular, are his illustrations of Judd's Margaret, and to Cooper's, Dickens's, and Simms's novels. In 1868, after some years' residence in Europe, he published both illustrations and letterpress of Sketches Abroad with Pen and Pencil. Died March 27, 1888.

Darling, a name applied to a river, a mountainrange, and two districts in Australia, is derived from Lieutenant-general Sir Ralph Darling (1775-1859), who was governor of New South Wales (1825-31). (1) The river Darling, a tributary of the Murray River, is formed by the union of several head-streams, all of which rise in the great Dividing Range. The chief of these head-streams are the Barwon and Gwydir. Farther down, the Darling receives the Culgoa and the Warrego from the north, and the Bogan from the south. Its length, with affluents, is stated at 1160 miles, and it and its tributaries are estimated to drain an area of 198,000 sq. m. It joins the Murray at Wentworth, on the border between New South Wales and South Australia, having received no tributary in its lower course. Much of the district traversed by it is an arid plain, save near the river-bank; the river is subject to floods. (2) The Darling Range, in Western Australia, runs parallel to the west coast, at a distance of 10 to 25 miles inland, from the southwest corner of the colony to a point some 250 miles farther north. The range has a height of from 1500 to 2000 feet; and in Mount William attains an elevation of 3000 feet. (3) The Darling district at the south-western corner of New South Wales, scantily watered, has an area of 50,000 sq. m. (4) The Darling Downs form the richest pastoral district of Queensland, in the southern part of the colony. This is an upland district on and about the summit of the Dividing Range. It was discovered by Allan Cunningham, the botanist, in 1827, and has an area of 6080 sq. m. There is also much fine agricultural land in which the cereals, potatoes, and all kinds of vegetables grow well. The western railway traverses the north, and the southern line the extreme east of this district.

Darling, GRACE, a name famous in the annals of heroism, was the daughter of William Darling (1795-1860), lighthouse-keeper on Longstone, one of the Farne Islands, and was born at Bamborough, 24th November 1815. On the morning of the 7th September 1838, the Forfarshire, bound from Hull to Dundee, with sixty-three persons on board, struck the Harker's Rock, among the Farne Islands, and in fifteen minutes forty-three persons were drowned. The vessel was seen by Grace Darling from the lighthouse at a quarter to five lying broken on the rocks. Darling and his daughter agreed that if they could get to her, some of the shipwrecked crew would be able to assist them in getting back. By wonderful strength and skill, they brought their boat to where the sufferers (nine in number) crouched. The solitary woman and four men were safely taken to the Longstone; two of the men returned with Darling, and succeeded in bringing the remainder

daring in itself, and so successfully carried out, filled every one with the warmest admiration. The lighthouse at Longstone, solitary and unknown no more, was visited by many of the wealthy and the great. Presents, testimonials, and money did not long survive her change of circumstances. were heaped at the feet of Grace Darling; but she She died of consumption, after a year's illness, on 20th October 1842. See Grace Darling, by E.

Hope (1876), and the Journal of William Darling

(1886).

Darlington, a parliamentary and municipal borough in the south of the county of Durham, on a slight elevation overlooking the Skerne near its junction with the Tees, 23 miles S. of Durham, and 45 NNW. of York. The chief industry of the place is connected with the extensive locomotive works, which give employment to many workers. There are iron and steel works in the north end of the town; brewing and tanning are carried on; and there are wool-mills. Pop. (1821) 6551; (1851) 11,228; (1871) 27,730; (1881) 35,102; (1891) 38,060, of whom many belong to the Society of Friends, long a prominent and influential element amongst the inhabitants. Darlington was created a parliamentary borough, sending one member to parliament, in 1867. Its prosperity dates from the opening, on 27th September 1825, of the Stockton and Darlington Railway, which was the first passengerline on which a locomotive-engine was employed; and that locomotive now stands on a pedestal outside the station. From the 11th century onwards the town belonged to the bishops of Durham, and till 1867 a borough bailiff, appointed by the bishop, managed its affairs; in that year it obtained a charter of incorporation. St Cuthbert's collegiate church, a very fine specimen of Early English, was founded in 1160 by Bishop Pudsey, and was restored by Sir G. G. Scott in 1869. It has three beautiful sedilia, and a tower 180 feet high. Among the

chief modern erections are the other churches

(Anglican, Presbyterian, Roman Catholic, and Nonconformist), the spacious new railway station (1887), a grammar-school, a high school for girls, a Britishi and Foreign School Society's college for female teachers, and the banks of Backhouse & Co., and Pease & Co. A free library was opened in 1885, for which Mr Edward Pease had bequeathed £10,000. At Oxen-le-field, 3 miles from Darlington, are curious cavities of unknown origin, called Hell Kettles; and near Darlington was the seat of George Allan the antiquary (1736-1800).

Darlingtonia, a genus of remarkable American Pitcher-plants, belonging to the family of Sarraceniacere (q.v.); and like its congeners, is insectivorous. See INSECTIVOROUS PLANTS.

Darmstadt, a town of Germany, capital of the grand-duchy of Hesse-Darmstadt, is situated on the small river Darm, 15 miles S. of Frankfort-on-Main. The streets of the old town are narrow, but those of the new town exhibit many imposing specimens of architecture. Darmstadt has several public squares, and fine public gardens and promenades. Besides the arsenal, the barracks, and the various religious edifices, it possesses two palaces. One of these, the old ducal palace, contains museums of painting, natural history, and archæology, and a library of 500,000 volumes; in the other, Prince Charles's palace, is Holbein's famous 'Meyer Madonna.' The handsome post-office dates from 1881, the theatre from 1871. There are manufactures of chemicals, hats, machinery, tobacco, playing-cards, carpets, and beer; and a trade in iron, petroleum, fruit, flour, and wine. But the place depends more on its ducal court and government offices than on

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its industries. Pop., with suburb (1875), 44,088;
(1885) 51,323; (1890) 56,503.
Darnel (Lolium temulentum), an annual grass
of the same genus with the valuable
Rye-grass (q.v.)-from which it may be
distinguished by the absence of the per-
ennial rootstock, by the empty glume,
equalling or exceeding the five to seven
(instead of eight to fifteen) flowered
spikelet, and the awn long instead of
short, or absent is common in corn-
fields in England and many parts of
Europe. It is the tares of the parable,
and has from ancient times been reputed
to have a narcotic poisonous seed, to
which many bad effects were ascribed,
which, in years of bad harvest, were
observed to ensue upon the eating of
bread or the feeding of horses upon oats.
The poisonous properties have been
denied by some botanists, and again
maintained by others, the most recent
investigations on the whole confirming
the ancient view. The seed may lie in
the ground for several years without
germinating, and then appear suddenly
in great quantity in a wet season.
It
can, however, be eaten by cattle with
impunity.

Darnel.

Darnétal, a town in the French department of Seine-Inférieure, 2 miles E. of Rouen, with manufactures of cotton, linen, and cloth goods. Pop. 6532. Darnley, HENRY STEWART, LORD, husband of Mary, Queen of Scots, was the eldest son of the Earl of Lennox by Lady Margaret Douglas, so a grandson of Margaret Tudor, and a cousin of Mary. Born in 1546, in England, where also he was educated, he was handsome in appearance, accomplished in manners, but fatally destitute of all moral and intellectual power. He perished by a gunpowder explosion at Kirk-of-Field, Edinburgh, 10th February 1567. Darnley is interesting chiefly on account of the position which he occupied with respect to his wife. See MARY, QUEEN OF SCOTS.

Darter (Plotus), a genus of birds nearly allied to Cormorants (q.v.), but very heron-like in gait and gesture. The four species are severally restricted to the warm parts of America, Africa,

Darter (Plotus anhinga).

Asia, and Australia. The head is small, with naked cheeks and throat; the neck is very long, thin, and flexible; the bill is long and straight; the tail and the toes are also long. The darters catch fish voraciously and with great dexterity; the name 'snake-neck' well describes them.

DARTMOOR

Though aquatic in habit, they rest and nest on trees. The American species figured is over three feet in length.

Dartford, a thriving market-town of Kent, in the narrow valley of the Darent, 2 miles above its influx to the Thames, and 17 ESE. of London. Edward III. here founded an Augustinian nunnery (1355); St Edmund's chantry was a great place of pilgrimage; and at Dartford Wat Tyler commenced his rebellion (1381). The church, with a Norman tower, was restored in 1863; among its interesting monuments is one to Sir John Spielman, Queen Elizabeth's jeweller, who in 1588 established at Dartford what is said to have been the first papermill in England. Paper is still manufactured, besides steam-engines and machinery, gunpowder, &c. The modern buildings include a county courthouse (1859), assembly rooms (1860), and, in the neighbourhood, the City of London lunatic asylum (1866). Pop. of parish (1851) 6224; (1881) 10,163; (1891) 11,962. See the local histories by Dunkin (1844) and Bayly (1876).

Dartmoor is a great granitic upland in Devonshire, the source, with two exceptions, of all the principal rivers of the county, remarkable alike for its wild and rugged scenery, its antiquities, its wide, solitary, trackless wastes, and its mineral products. It is upwards of 130,000 acres in extent, the extreme

length from north to south being 25 miles, and the

extreme breadth from east to west 20 miles. The outline is irregular. The central portion is the ancient royal forest of Dartmoor, and this is surrounded by a belt of open country, once known as the Commons of Devonshire,' but portions of which have been inclosed. The attempts to cultivate Dartmoor itself have been very few, and the northern quarter for miles shows no trace of man, but the valleys through which the rivers descend to the lowland country are singularly fertile, and at times full of beauty. The moor itself affords valuable mountain pasture to cattle, sheep, and large numbers of halfwild ponies. The forest rights belong to the duchy of Cornwall; but there are rights of pasturage exercised by holders of what are called venville tenures in certain parishes bounding the moor, which date prior to the Norman Conquest.

The average height of Dartmoor above the sea is upwards of 1200 feet, but its highest point, High Willhayes, is 2039 feet; and the next, Yes Tor, 2030. The hills are commonly called tors, and for the most part have granite crests, weathered into grotesque and picturesque shapes. The whole of Dartmoor is of granite, protruded between the close of the Carboniferous and the opening of the Triassic periods. Devonian rocks mantle round its southern extension, and Carboniferous round its northern, associated at various points with gabbros, dolerites, and other intrusive rocks. The granite is chiefly gray, but there are rich red varieties, as at Trowlesworthy. Large quantities of the ordinary stone have been quarried, especially at Hey Tor and King Tor. Dartmoor is rich in minerals. Tin has been raised for many centuries, long before the dawn of history, by streaming in the valleys, and vestiges of the ancient mining operations of the old men' abound. Copper, iron, and manganese have also been worked, but mining is now carried on at a few points only. Gold has frequently been found in the river-beds. The tinners of Devon had a quasi-corporate existence in Saxon times, and their rights were confirmed by King John and other monarchs. The Stannary Parliaments, in which they managed their affairs, were held in the open air on Crockern Tor. The most important mineral product of Dartmoor at the present day is china-clay, or kaolin, which is the result of the decomposition of the felspar

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DARTMOUTH

of the granite. The largest china-clay works in England are at Lee Moor, and are connected by a tramway with wharves at Plymouth.

The fauna and flora of Dartmoor have many points of interest. Trees are very rare; but there is a very singular group of gnarled and stunted oaks on a slope overlooking the West Dart, near Crockern Tor, of high antiquity and weird aspect, which has been called one of the wonders of the moor' "Wistman's Wood.'

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The chief rivers rise at a height of over 1800 feet above the sea, in a wide stretch of peat-bog around Cranmere Pool-the Dart, Teign, Taw, Okement, Lyd, Tavy, and Walkham. From the morasses of the southern quarter spring the Plym, Yealm, Erme, and Avon.

Dartmoor is unrivalled in England in the extent and character of its prehistoric and rude stone antiquities-earthworks, barrows, kistvaens, menhirs, lines or avenues, cyclopean bridges, circles, cromlechs, trackways, and pounds or inclosures of stones, sometimes containing the remains of villages. Many stone implements have been found. On several of the tors there are rock basins, formerly called Druidical, but now assigned to the operation of natural causes.

The chief centre of population is Prince Town, named after George IV. when Prince Regent. Here a prison was built (1806) during the war with Napoleon, for the reception of prisoners of war. When the war ended it was abandoned, and was at one time used for the manufacture of naphtha from peat, which failed, as all attempts to utilise the Dartmoor peat beyond the supply of local wants have done. In 1855 the buildings were adapted to their present purpose of a convict prison. Attached is a fertile reclaimed farm. A very picturesque railway runs to Prince Town. See the Rev. S. Rowe's Perambulation of Dartmoor (1856).

Dartmouth, a seaport and municipal borough (till 1867 also parliamentary) in the south of Devonshire, 32 miles S. by W. of Exeter. It is built in picturesque terraces on a steep slope 300 to 400 feet high, on the right bank of the romantic estuary of the river Dart, at a short distance from the sea. The streets are narrow, and many of the houses very old, with overhanging stories, projecting gables, and wood-carvings. St Saviour's Church (circa 1372) has a richly sculptured, painted, and gilt stone pulpit, and a beautifully carved rood-loft. A battery, and the remains of a castle built during the reign of Henry VII., stand at the entrance to the harbour. Pop. (1891) 6038. Dartmouth is a quarantine port of the Eng lish Channel, and has a considerable trade with the Mediterranean, and in the Newfoundland fisheries. It is the port of departure of the 'Castle' line of mail steamers to South Africa, and a favourite yachting station—the harbour being deep and landlocked. At Dartmouth, in 1190, the Crusaders, under Richard Coeur-de-Lion, embarked for the Holy Land. The French burned it in Richard's reign, but were repulsed, chiefly by the women, in an attack in 1404. In 1346 Dartmouth furnished 31 ships for the siege of Calais. In 1643 Prince Maurice besieged and garrisoned it; but in 1646 Fairfax stormed and took it. Newcomen, the inventor of the steam-engine, was born here; Sir Humphrey Gilbert was born at Greenway, on the opposite side of the Dart; and another great Elizabethan worthy, John Davis, at Sandridge.

Daru, PIERRE ANTOINE, COMTE, a French writer and financier, was born at Montpellier, 12th January 1767. At sixteen he entered the army, and rose rapidly, but was flung into prison during the Terror, where he amused himself by translating Horace. Under Napoleon he was Intendant

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general in Austria and Prussia, and a councillor of state, while in 1818 Louis XVIII. made him a peer. Thenceforth he devoted himself exclusively to letters. He died a member of the Institute and of the Academy of Sciences, 5th September 1829. Of his many books the chief are Cléopédie, ou Théorie des Réputations Littéraires (1800), spirited poem; Histoire de la République de Venise (7 vols. 1819-1821); and Histoire de la Bretagne (3 vols. 1826).-His son, NAPOLÉON DARU, born in 1807, opposed the coup d'état, and was proscribed; but became a member of the National Assembly in 1871, of the senate in 1876. Died in 1890.

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Darwen, a municipal borough of Lancashire, on the river Darwen, 3 miles S. of Blackburn, and 9 N. of Bolton. Cotton is the staple manufacture; then come paper-making and paper-staining; and to these and other industries, with its water facilities, and the neighbouring coal-mines and stone quarries, Darwen owes its rapid growth and its well-being. It was incorporated in 1878. Among the chief buildings are the free library, the market erected in memory of Sir Robert Peel. Pop. (1851) hall, the co-operative hall, and the public baths, 7020; (1881) 29,744; (1891) 34,192.

Darwin, CHARLES ROBERT, F.R.S., naturalist, the discoverer of the principle of natural selection, was born at Shrewsbury, February 12, 1809. His grandfather was Dr Erasmus Darwin, and his father Dr Robert W. Darwin, F.R.S. His mother was a daughter of Josiah Wedgwood, the celebrated potter. Darwin was educated at Shrewsbury grammarschool, studied at Edinburgh University for two sessions in 1825-6-7, and entered at Christ's College, Cambridge, in 1828. Already at Edinburgh he had become a member of the local Plinian Society; he took part in its natural history excursions, and read before it his first scientific paper-a new contribuIt was at Cambridge, however, that his biological tion to our knowledge of the Flustra or sea-mats. studies seriously began. Here he became acquainted with Professor Henslow, the well-known botanist, who encouraged his interest in botany and zoology. His chief taste at this time was for geological research. In 1831 he took his degree of B.A., and shortly after was recommended by Beagle, under Captain (afterwards Admiral) FitzHenslow as naturalist to the expedition of H.M.S. roy, R.N., then about to start for a scientific survey of South American waters. He sailed on December 27, 1831, and did not return to England from his long cruise till October 2, 1836. Meanwhile he visited Teneriffe, the Cape de Verde Islands, the Brazilian coast, Monte Video, Tierra del Fuego, Buenos Ayres, Valparaiso, and the New Zealand, Tasmania, and the Keeling Islands, Chilian region, the Galapagos Archipelago, Tahiti, in which last he laid the foundation for his famous theory of coral reefs. It was during this long expedition that Darwin obtained that intimate knowledge of the fauna, flora, and geological conditions of many tropical, subtropical, and temperate climates which so admirably equipped him at last for the great task he was afterwards to perform in settling the factors of biological evolution.

On his return to England in 1836, he set to work to co-ordinate the results obtained during his voyage. other scientific leaders, by whose aid he was apHe formed the friendship of Sir Charles Lyell and pointed secretary of the Geological Society in 1838. A year later, he was elected to the fellowship of the Royal Society, and early in 1839 he married his cousin, Miss Wedgwood. In the same year he published his Journal of Researches into the Geology and Natural History of the various Countries visited by H.M.S. Beagle. From 1840 to 1843 Darwin was occupied with the publication of the Zoology of the

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Voyage of the Beagle, under government auspices, to which great work (compiled by the leading specialist authorities of the day) he himself contributed an introduction and many notes. In 1842 appeared his work on The Structure and Distribution of Coral Reefs; in 1844, Geological Observations Volcanic Islands; and in 1846, Geological Observations on South America. These works placed him at once in the front rank of contemporary scientific thinkers. In 1851-53 appeared his valuable treatise on barnacles, A Monograph of the Cirripedia.

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Three years after his marriage Darwin settled at Down, near Beckenham, in Kent, where for the rest of his days he passed his time as a country gentleman among his conservatories, his pigeons, his garden, and his fowls. The practical information thus gained (especially as regards variation and interbreeding) was of invaluable use to him in his later researches. Private means enabled him to devote himself unremittingly henceforth, in spite of continuous and distressing ill-health, to the pursuit of science. It was at Down that Darwin began to occupy himself with the great work of his life-the problem of the origin of species. In 1837 he had already set out to accumulate facts and observations for this purpose. After five years' unremitting work, he allowed himself to speculate' on the subject, and drew up some short notes, which he enlarged in 1844 into a sketch of conclusions for his own use. These conclusions embodied in embryo the famous principle of natural selection, the germ of the celebrated Darwinian Theory (q.v.). With constitutional caution, however, Darwin delayed publication of his hypothesis, which was only at length precipitated many years later by an accidental circumstance of a romantic character. In 1858 Mr Alfred Russell Wallace, the distinguished explorer, sent home from the Malay Archipelago a memoir addressed to Darwin himself, for presentation to the Linnean Society. On opening this packet, Darwin found to his surprise that it contained in essence the main idea of his own theory of natural selection. Sir Charles Lyell and Sir Joseph Hooker, to whom he communicated the facts, persuaded Darwin to read a paper of his own concomitantly with Wallace's before the Linnean Society, which was accordingly done on July 1, 1858. Urged forward by this strange coincidence, Darwin set to work seriously at once to condense his vast mass of notes-the labour of a lifetime and put into shape his great work on The Origin of Species by means of Natural Selection, published in November 1859. For an account of the main ideas there promulgated, see DARWINIAN THEORY. The book itself, an epoch-making work, was received throughout Europe with the deepest interest, was violently attacked and energetically defended, but in the end succeeded in obtaining recognition (with or without certain reservations) from almost all competent biologists. From the day of its publication Darwin continued to work on unremittingly at a great series of supplemental treatises, in which his main principles were still further enforced and enlarged, while minor corollaries were brought prominently into view. The Fertilisation of Orchids appeared in 1862, The Variation of Plants and Animals under Domestication in 1867, and The Descent of Man in 1871. The lastnamed work, hardly less famous than the Origin of Species, derives the human race from a hairy quadrumanous animal belonging to the great anthropoid group, and related to the progenitors of the orangutan, chimpanzee, and gorilla. In this book Darwin also developed his important supplementary theory of sexual selection, which on the whole has been received by scientific thinkers with less favour than his other ideas. His later works are The

Expression of the Emotions in Man and Animals (1873), Insectivorous Plants (1875), Climbing Plants (1875), The Effects of Cross and Self Fertilisation in the Vegetable Kingdom (1876), Different Forms of Flowers in Plants of the same Species (1877), and The Power of Movement in Plants (1880). His last work, The Formation of Vegetable Mould through the action of Worms, appeared in 1881. In it Darwin showed grounds for believing that the vegetable mould which covers a large part of the globe is mainly due to the castings of earth-worms, without which the greater portion of the land surface of the world must necessarily consist of barren rock or thirsty sand.

It is, however, as the great leader of evolutionary biology that Darwin will be mainly remembered among men. Though not (as is commonly, but erroneously, believed) himself the originator of the evolution hypothesis, nor even the first to apply the conception of descent with modification to plant and animal organisms, Darwin was undoubtedly the first thinker to gain for that conception a wide acceptance among biological experts. By adding to the crude evolutionism of Erasmus Darwin, Lamarck, and others, his own specific idea of natural selection, he supplied to the idea a sufficient cause, which raised it at once from the level of a hypothesis to the grade of a verifiable theory. His kindliness of character, honesty of purpose, devotion to truth, and attachment to his friends, rendered him no less remarkable on the moral and emotional than on the purely intellectual side of his nature. For many years his health had been extremely feeble, and he had worked under the severest physical disadvantages. He died suddenly, after a very short illness, April 19, 1882, and was buried with unusual honours in Westminster Abbey. See his Life and Letters by his son, Francis Darwin (1887).

Darwin, ERASMUS, physician, natural philosopher, and didactic poet, was born 12th December 1731, at Elston Hall, near Newark, in Nottinghamshire; entered St John's College, Cambridge, in 1750, graduated B.A. in 1754, and afterwards studied medicine at Edinburgh. After an unsuccessful attempt to establish a practice at Nottingham, he removed to Lichfield, where he married and became a popular physician and prominent figure from his ability, his radical and free-thinking opinions, his poetry, his eight-acre botanical garden, and his imperious advocacy of temperance in drinking. After his second marriage in 1781, he settled in Derby, and then at Breadsall Priory, where he died suddenly 18th April 1802. Darwin had once a great reputation as a physiologist, but his system is, for the most part, inconsequential, baseless, and untenable. At the same time, many of his ideas are original, suggestive, and contain within them the germs of important truths. His strength and his weakness lay in his faculty for seeing analogies in nature. Sometimes he is exceedingly happy in his discoveries; at other times he is quite fantastical. The same remarks hold good as regards his verse, where, amid the frequent extravagance and incomprehensibility of his notions, there burst forth strains of genuine poetry. The Loves of the Plants (1789), a part of his chief poem, the Botanic Garden, was happily burlesqued in the 'Loves of the Triangles' in the Anti-Jacobin. Interest in Darwin's speculations has been revived by the recognition of his partial anticipation of Lamarck's views on evolution, and so of his own famous grandson's. His chief works are Zoonomia, or the Laws of Organic Life (1794-96); and his Phytologia, or Philosophy of Agriculture (1799). See his Life by Krause, trans. by Dallas (1879). By his first wife he was grandfather of Charles Darwin; by his second, of Francis Galton.

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DARWINIAN THEORY

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Darwinian Theory, the explanation of the species, specially concentrating himself upon what evolution of forms of life on our planet, offered seemed to him the weakest point of the preceding by the great naturalist, Charles Darwin (q.v.), theories, the explanation of adaptations. ComFrom the very outset we must carefully guard mencing in 1837, after five years' work, I allowed against the confusion, still widely popular, of myself to speculate on the subject, and drew up 'Evolution' with 'Darwinism.' Evolution must some short notes; these I enlarged in 1844 into a clearly be retained to denote the entire drama of sketch of the conclusions which then seemed to be cosmic change; Darwinism, therefore, must as probable; from that period to the present day I clearly be restricted to one particular interpreta- have steadily pursued the same object.' This was tion of the mechanism and plot of this cosmic written as late as 1859, and even then because he drama, of many which have been thrown out by had received a paper from Mr A. R. Wallace (at reflective spectators. Darwin expressed the mode that time exploring the Malay Archipelago), in of the evolutionary process in a classic phrase, which views identical with his own were expressed, the title of his great work-The Origin of Species and thus was compelled to proceed to the publicaby means of Natural Selection, or the Preservation tion of his results. He did so first in a brief outof Favoured Races in the Struggle for Life. That line read, by advice of Lyell and Hooker, simulexplanation is not only in itself peculiarly shrewd taneously with Wallace's paper (see Jour. Linn. and well reasoned, luminous and widely appli- Soc., 1858); and in the following year in that fuller cable, but has been of the very greatest import- abstract, as at first he merely regarded it, which ance in awakening biologists and students of all soon became so famous-the Origin of Species. other sciences, and thus ultimately the intelligent Its substance may with advantage be briefly sumpublic, to the facts of the case-to the conscious- marised, so far as further compression of such ness of this evolutionary drama. On the pro-intellectual pemmican' is possible. For details mulgation of this new hypothesis as to the mode of and explanations the reader must consult the work occurrence of evolutionary changes in the organic itself (7th ed. 1880). world, the whole discussion as to the occurrence of evolutionary change at all speedily came to centre round it; although in view of the powerful arguments for the occurrence of evolution which had been independently adduced by Spencer and others, it may be admitted this was not quite logical. The fact, however, remains. Moreover to the new standpoint afforded by the clear acceptance of both ideas -evolution and this through natural selection-and to the energetic application of these, first by Darwin himself, but soon by a multitude of zealous workers, we owe a progress which it would as yet be premature to estimate, but which has pervaded the whole field of biology, and even the fields of all the higher sciences, mental and social, which so largely utilise biological methods and generalisations. Whatever may be the subsequent development of our evolutionary conceptions, the epoch-making importance of the Darwinian theory will be unaltered. Hence the expediency of the present comparatively full exposition of its main positions and of their bearings, apart from the larger and more general argument under EVOLUTION.

To the statement of Darwin's theory, therefore, we inay at once proceed, postulating no more than that general acquaintance with the aims and results of biology which is now becoming so commonly current, or which may readily be gained by help of the articles BIOLOGY and BOTANY. Nor need any statement of the general doctrine of evolution, or of evolutionary theories before Darwin, here detain us, since these find more fitting place in the general article EVOLUTION.

The failure of pre-Darwinian theories to gain any very general acceptance among naturalists was no doubt largely attributable to established prejudice, backed as this was by the predominant authority of Cuvier. This was by no means the whole explanation. For while those theories rendered it extremely probable that modification had occurred, they all fell short, as Darwin pointed out, in one most important particular. They failed to show how the modification of one species from another could take place, so as to acquire that perfection of structure and co-adaptation which justly excites our admiration;' since the hypotheses of the potency of external conditions, of habit, of the volition of the organism itself, and so on, alike successively broke down.

Darwin was especially struck by the distributional phenomena he witnessed during his Naturalist's Voyage,' and thereafter devoted himself primarily to the problem of the origin of

Outline of Origin of Species.'-In order to gain insight, then, into the means of modification, Darwin commences with a study of the variation of plants and animals under domestication (later expanded into a separate work; 2d ed. 1876). Variation and Heredity.-While all plants and animals exhibit some degree of variation, this is greatest among domesticated species, owing to their new and less uniform conditions of life. These may act directly on the whole organisation, or on separate parts, and the variation, though rarely, is sometimes definite, as when size increases with quantity of food, or colour changes with its quality; or the conditions may act indirectly by influencing the reproductive system, which is peculiarly sensitive. Changed habits produce an inherited effect-e.g. the leg-bones of the common duck weigh proportionally more, and its wing-bones less, than in the wild variety, because it flies less and walks more. So, too, tame mammals acquire drooping ears, since these are rarely pricked in alarm. One variation is usually correlated with others, thus long-beaked pigeons have small feet, and conversely. All variations tend to be inherited. The popular belief that domestic races simply revert to the aboriginal stock is unsupported by facts.

Save that domestic varieties are less uniform than wild species, often differ more widely in some single part, and are fertile when crossed, there is no well-marked distinction between these and socalled true species. If, therefore, such varieties as the different breeds of the dog can be shown to be descended from a single wild species, there necessarily arises great doubt as to the immutability of closely allied natural species, such as the foxes. While, however, the many breeds of dog appear to have arisen from several wild species, and those of cattle also from two or three, our fowls, ducks, rabbits, &c., all certainly arise from a single ancestral species. The case of pigeons is of peculiar importance, since pouter, carrier, fantail, and tumbler differ so thoroughly, externally and internally, that any ornithologist would be compelled to assign to them not merely specific but generic distinctness, if he had discovered them in the wild state. is at least as much difficulty in believing that such breeds can have proceeded from a common ancestor, as there is in the case of any group of birds in nature; and every breeder of these has been firmly convinced of their descent from distinct species. Yet all these breeds are proved to come from the common rock-dove (Columba livia; see PIGEON),

There

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